have subsisted
Present perfect conjugation of subsist. There are other translations for this conjugation.

subsist

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It has been asked by the opponents of such views as I hold, how, for instance, could a land carnivorous animal have been converted into one with aquatic habits; for how could the animal in its transitional state have subsisted?
Los adversarios de las ideas que sostengo han preguntado cómo pudo, por ejemplo, un animal carnívoro terrestre convertirse en un animal con costumbres acuáticas; porque ¿cómo pudo subsistir el animal de su estado transitorio?
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